Western Pond Turtle Literature – Citations New and Old

For those of you western pond turtle aficionados, I hope by now you have stumbled across the western pond turtle literature page, where the bulk of the western pond turtle literature published between 1852 and the present has been meticulously indexed and posted for public consumption. I say ‘bulk’ because even I overlook a paper now and then, or fall behind in my indexing. But thanks to the gentle prodding of GL and LP – two fellow western pond turtle wonks – I’ve picked up where I last left off and offer the following new (and old) publications in case you too have overlooked them.

Abel, Jae. 2010. Western Pond Turtle Summer Habitat Use in a Coastal Watershed. Unpublished M.S. Thesis: San Jose State University. 111 pp.

Angielczyk, Kenneth D., Chris D Feldman, and Gretchen R. Miller. 2010. Adaptive Evolution of Plastron Shape in Emydine Turtles. Evolution. 65(2): pp 377-394.

Ashton, Don T., Jamie B. Bettaso, and Hartwell H. Welsh Jr. 2011. Comparative Ecology of Western Pond Turtle (Actinemys marmorata) Populations on the Free-Flowing South Fork and Regulated Main Fork Trinity River: Demography, Size and Body Condition Comparisons, Thermal Ecology, and Spatial Dynamics. Final Report to the Trinity River Restoration Program. June 10. 55 pp.

Bettelheim, Matthew P., R. Bruce Bury, Laura C. Patterson, and Glen M. Lubcke. 2006. Trachemys scripta elegans (Red-eared Slider). Reproduction. Herpetological Review 37(4): pp 459-460.

Bettelheim, Matthew P. 2009. Actinemys (=Clemmys) marmorata (Western Pond Turtle). Courtship Behavior. Herpetological Review 40(2): pp 212-213.

Bettelheim, Matthew. 2010. Native Son: Making Room for the Western Pond Turtle. Bay Nature. 10(2): April-June pp 40-44.

Bettelheim, Matthew P. 2011. Art in Herpetology. Herpetological Review 42(3): p 382.

Bettelheim, Matthew. 2011. Western Pond Turtles: Basking in Their Own Resiliency. Outdoor California. 72(2): March-April pp 22-31.

Bury, R. Bruce, David J. Germano, and Gwendolynn W. Bury. 2010. Population Structure and Growth of the Turtle Actinemys marmorata from the Klamath-Siskiyou Ecoregion: Age, Not Size, Matters. Copeia. 3: pp 443-451.

Bury, R. Bruce, Hartwell H. Welsh, David J. Germano, and Donald T. Ashton. 2012. Western Pond Turtle: Biology, Sampling Techniques, Inventory and Monitoring, Conservation and Management. Northwest Fauna 7: 128 pp.

Fritz, Uwe and Peter Havas. 2007. Checklist of Chelonians of the World. Vertebrate Zoology. 57(2): pp 149-368.

Fritz, Uwe, Christian Schmidt, and Carl H. Ernst. 2011. Competing Generic Concepts for Blanding’s, Pacific and European Pond Turtles (Emydoidea, Actinemys and Emys)–Which is Best? Zootaxa. 2791: pp 41-53.

Germano, David J. and R. Bruce Bury. 2009. Variation in Body Size, Growth, and Population Structure of Actinemys marmorata from Lentic and Lotic Habitats in Southern Oregon. Journal of Herpetology. 43(3): pp 510-520.

Germano, David J. 2010. Ecology of Western Pond Turtles (Actinemys marmorata) at Sewage-Treatment Facilities in the San Joaquin Valley, California. The Southwestern Naturalist. 55(1): pp 89-97.

Gordon, Rebecca A. 2009. Effects of Incubation Temperature on the Embryonic Development and Hatching Success of the Western Pond Turtle (Emys marmorata). Unpublished M.S. Thesis: Sonoma State University. 38 pp.

Lambert, M.R., S.N. Nielsen, A.N. Wright, R.C. Thomson, and H.B. Shaffer. In press. Microhabitat Characteristics Favor Introduced Red-Eared Sliders Over Threatened Western Pond Turtles in a Human-Dominated Landscape. Chelonian Conservation and Biology.

Lightfoot, Kent G. and Otis Parrish. 2009. California Indians and their Environment, An Introduction. California Natural History Guide Series No. 96. University of California Press: Berkeley, California. 512 pp.

Polo-Cavia, N., T. Engstrom, P. Lopez, and J. Martin. 2010. Body Condition Does Not Predict Immunocompetence of Western Pond Turtles in Altered Versus Natural Habitats. Animal Conservation. 13(3): pp 256-264.

Reynolds, Robert P., Steve W. Gotte, and Carl H. Ernst. 2007. Catalog of Type Specimens of Recent Crocodilia and Testudines in the National Mueum of Natural History, Smithsonian Institution. Smithsonian Contributions to Zoology Number 626. 49 pp.

Rosenberg, Daniel K. and Roberta Swift. 2010. Post-Emergence Behavior of Hatchling Western Pond Turtles. Final Report. Oregon Wildlife Institute, Corvallis, Oregon. 33 pp.

Shufeldt, R.W. 1919. Observations on the Chelonians of North America (Part IV). Aquatic Life. 4(12): pp 155-157.

Spinks, P. Q., R. C. Thomson, and H. B. Shaffer. 2010. Nuclear Gene Phylogeography Reveals the Historical Legacy of an Ancient Inland Sea on Lineages of the Western Pond Turtle, Emys marmorata in California. Molecular Ecology 19: pp 542-556.

Sproat, Gilbert Malcolm. 1868. Scenes and Studies of Savage Life. Smith, Elder and Co.: London. 317 pp.

Stebbins, Robert C. and Samuel M. McGinnis. 2012. Field Guide to Amphibians and Reptiles of California (Revised Edition). University of California Press: Berkeley, California. 552 pp.

Studebaker, Rebecca S. 2008. Actinemys marmorata (Western Pond Turtle). Predation Attempt. Herpetological Review. 39(4): pp 463-464.

Thompson, Robert C., Phillip Q. Spinks, and H. Bradley Shaffer. 2010. Distribution and Abundance of Invasive Red-Eared Sliders (Trachemys scripta elegans) in California’s Sacramento River Basin and Possible Impacts on Native Western Pond Turtles (Emys marmorata). Chelonian Conservation and Biology. 9(2): 297-302.

White, W.L. 1910. Turtles and Turtling in Many Waters. Hunter Trader Trapper. 20(2): pp 17-20.

Wilcox, Jeffery T. 2010. Actinemys (=Emys) marmorata (Western Pond Turtle). Predation. Herpetological Review. 41(2): pp 212.

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  1. #1 by Don Stephens, life-long student of 'Clemmys' on April 26, 2013 - 11:08 am

    Wow! Matt!

    What a wonderful resource you and your blog/website are for us lay-students of marmoratas! And your up-date is super…

    I just wish more of these academic papers had links to them or to their authors’ vita sites / email e-ddresses! How might a lay-person seek out copies of the ones published by scientific journals? Any suggestions?

    It would also be great if you could post your papers and articles to your site… : )

    Appreciating your blog and its sharing more each day!

    – Don Stephens,

  2. #2 by Matthew Bettelheim on April 26, 2013 - 1:17 pm

    Thanks as always, Don.

    The beauty and bane of the internet is the propensity for links to change over time, so while I often link to digital sources, it’s hard to stay abreast as sites that host these papers evolve.

    The best way to find a paper you’re interested in is to query the article title together with “pdf”; you’ll be amazed at how many papers have been digitized and are available online. Even better is to visit an author/researcher’s professional website. They often post papers that are otherwise “locked up” behind a pay-wall or journal subscription.

    As for my pubs, you make a good point. It may take some time, but let me contact the individual journals for the necessary permissions. I’ll see what I can do.


  3. #3 by Don Stephens, life-long student of 'Clemmys' on April 27, 2013 - 4:15 pm

    Matt, Visiting Jeffrey Lovich’s professional webpage:


    ….is how I found current access to his papers, since the previous direct gov-links for them are now corrupted…There must be over 70 linked papers there, including a number regarding PPTs, a fair number of which don’t have him as the lead author, so might otherwise be difficult to trace.

    Thanks, again,Matt….Please keep up your good work

    – Don Stephens

  4. #4 by Don Stephens, life-long student of 'Clemmys' on May 8, 2013 - 1:29 am

    Hi again, Matt,

    I notice that your listing above, for your own excellent and beautifully photo-illustrated article:

    Bettelheim, Matthew. 2011. “Western Pond Turtles: Basking in Their Own Resiliency”, Outdoor California. 72(2): March-April pp 22-31.

    …does not include the publicly-posted, California Department of Fish and Wildlife link for it…..you might want to add that, when you integrate your two PPT Liturature Bibliographies, (which I presume you are in the process of doing now.) It’s at:


    ….In case you don’t have it at hand, currently… : )

    – Don Stephens

  5. #5 by Matthew Bettelheim on May 13, 2013 - 6:32 pm

    Thanks Don, I’ve added the link to the post. Good suggestion. Updates to the biblio to follow shortly…

  1. Western Pond Turtle Literature – Citations New and Old | (bio)accumulation

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